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Fig. 3. The movement of an acentric fragment generated by cutting a trailing arm
from a segregating half-bivalent conformed to one of three types. (A-C) The
single trailing arm that was to be severed is represented by an open rectangle
and the acentric fragment generated by cutting is represented by an open
square. The kinetochore-containing fragment is represented by an open square
connected to a circle, the latter representing the centromere/kinetochore
domain of the half-bivalent. For simplicity, only the to-be-cut arm and its
partner are included. (A) Type 1 fragment: the fragment moved rapidly
backwards across the equator into the opposite half spindle (A2) to make
contact with its partner (A3) and then moved along with its partner to the
lower pole (A4). The tethered arm of the partner half-bivalent is shown with a
solid rectangle and solid circle. (B) Type 2 fragment: the fragment moved
backwards across the spindle into the opposite half spindle (B2), but it did
not make contact with its partner (B3) as in (A). Further movement of the
fragment to the opposite pole occurs after the partner (solid) reaches the
pole (B4). (C) Type 3 fragment: this fragment moved backwards a short distance
(or did not move backwards at all) without crossing the equator (C2) then
shifted from backward movement to movement to the correct pole (C3) and at
late anaphase it was located near its original pole (C4).