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Fig. 5. Viscoelastic properties of the intranuclear region and the cytoplasm. (A) Mean frequency-dependent elastic [G'({omega})] and viscous [G''({omega})] moduli of the intranuclear region (black lines) and cytoplasm (red lines). Open symbols: G'; closed symbols: G''. (Inset) Frequency-dependent phase-angle for cytoplasm (black line) and intranuclear region (red line), {delta}({omega})=tan—1[G'' ({omega})/G'({omega})]. Examplary angles for liquids (e.g., glycerol), {delta}=90° and elastic solids (e.g. concrete), {delta}=0°. Mean moduli G' ({omega}) and G'' ({omega}) were calculated from the ensemble-averaged MSD, , as described (Mason et al., 1997). Distribution of (B) elastic and (C) viscous moduli, G' and G'', respectively, in the intranuclear region measured at a frequency of 1 Hz. (D) Distribution of shear viscosity {eta}s [which is approximately the product of the plateau modulus G'p (inset in B)] and the relaxation time {tau}R (inset), {eta}s=G'p{tau}R (see text for details). For panels B-D, n=84; 1 Poise (P) = 1 Pascal second (Pa.s); 1 dyn/cm2 = 0.1 Pa = 0.1 N/m2.





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